The reference state for semi-natural neutral grassland is species-rich grassland, hay meadow or pasture, where the cover of herbs is greater than the cover of grasses. This reference state described here occurs on soils leaning towards the more acid end of the neutral spectrum. In Wales, these meadows, which can support populations of wood bitter vetch (Vicia orobus), greater burnet (Sanguisorba officinalis) and wood anemone (Anemone nemerosa), are typically, though not exclusively, found at more elevated locations near the upland fringe.